Ground colór of alcohol-preserved specimens light straw color. Body crossed by four wide bars subject to considerable variatíon in intensity; dark coloration of bars produced by black scale margins to give a crosshatched appearance. Fírst bar originating under fírst spine of second dorsal fin, but on sorne indivi duals may be continuous with darkened nape. Second bar under posterior half of second dorsal fin and may almost merge with fírst bar. Third bar under anterior half of third dorsal fin. Fourth bar on caudal peduncle; most intense in males. Only last two bars usually reaching ventr al midline. Several dark markings on head: a bar at midposterior border of orbít, an oblique bar from eye to lower margin of cheek, an oblique bar between eye and expanding on upper lip. Posterior border of preopercle black;
J. barbarensis and J. costacubensis, twospecies present along the Pacific and Caribbean coasts ofCostaRica, respectively, differ from J. anulatus. J. barbarensis has a translucent white body with gold and blue cerata, mark- ings on the dorsal papillae with branched digestive gland ducts, jaws with denticles, and lateral teeth with denticles. J. costacubensis has a cream body with a longitudinal white line, rhinophores with papillae on the bases, and lamellae in the distal part of the rhinophores. In this species, the entire epithelium of the dorsal papillae bears papillae, while in J. anulatus only the apical portion above the ring bears papillae. The digestive gland in the papillae of J. costacubensis extends for more of the length than does that of J. anulatus.
Elytra l.20-1.36 (1.28) times longer than wide, 1.48-1.62 (1.52) longer than pronotum; sides almost straight and parallel on basal two-thirds, rather narrowly rounded behind; striae not impressed, punctures small, shallow, spaced by less than diameter of a puncture; interstriae two times wider than striae, punctures two-third width of strial punctures, mostly uniseriate; interstriae 10 acutely elevated to sternum 3; declivity rather steep, convex; strial and interstrial punctures as on disco Vestíture consisting of interstrial rows of spatulate bristles, mostly on declivity, bristles as long as distance between rows, slightly closer to one another within a row; minute hairlike setae in rows on striae and interstriae.
montré l' existence d' un couloir de distribution. avec des formes en gradient... Notre premiere réaction á l' égard du matériel de CostaRica a été de proposer une nouvelle espéce ..... Seule. l'étude d'un matériel beaucoup plus important, associé a l'étude de son écologie, pourra montrer l'existence ou non de véritables formes indépendantes." In their monograph on the scorpions ofCostaRica, Francke and Stockwell (1987) accepted my 1982 decision and redescribed the Ananteris fromCostaRica
about 12 papillae with a longitudinal branch of five papillae posteriorly and another of three papillae below posteroventral border of eye. Three longitudinal rows: a row of four papillae behind angle of mouth; a double curving row on the lower preopercular arm, 19 small papil lae with a parallel row of five widely-spaced larger papillae below. Postorbital series - An oblique row of four papillae on nape behind posterodorsal margin of eye. Interorbital series (not visible on lateral view) - Six papillae crossing the mid interorbital space; one isolated papilla on anterodorsal margin of eye and another on posterodorsal margin of eye. Preorbital series - An oblique row of 15 papil lae on side of snout below posterior nostril, curving down to maxilJa; two larger isolated papillae closely anterior to oblique row. Mandibular series - A double series of a row of 11 small papillae aboye a parallel row of five larger papillae; a longitudinal row of five papil lae on each side of isthmus (not visible on lat eral view of Fig. 2). Oculoscapular series - Five papillary tracts of di verse orientation along oculoscapular sulcus; an oblique row of 6 papillae immediately below sulcus near pos terodorsal comer of preopercle. Opercular series - An oblique curved row of five papillae on mid-opercle and a transverse row of 21 papilJae near anterior margin of opercle.
The basal color of Undulambia adults is brown, not white, as in Albusambia and Neurophyseta. Wings of Undulambia are incised, and those of Albusambia and Neurophyseta are entire. Forewing costal swelling is absent in Albusambia and Neurophyseta, but pres- ent in Undulambia. In the male genitalia of Undulambia the valva is long and equal in width throughout, not widened posteriorly as in Albusambia and some Neurophyseta spe- cies. The body of Undulambia larva is round, not dorsoventrally flattened, and not inter- segmentally constricted as in Albusambia. In Undulambia the mandible consists of one line of teeth, not two as in that of Albusambia. Other than the host plant record, the bio- logical and morphological data of the larvae of Neurophyseta are not available. Unlike the vertex of the pupal head of Albusambia, Undulambia does not have a prominent medial dorsoventral depression. Undulambia has a somewhat rugose, broadly flattened prothorax with two anterolateral horn-like structures that protrude twice as much as in the smooth pro- thorax of Albusambia. Albusambia has lateral conical depressions posterior to the anus, but Undulambia lacks these depressions. Pupal data on Neurophyseta is incomplete.
Colomb:a 900 miles southeast, and northern Venezuela 1000 miles east of the Costa Rican locale. Although closely allied to other American species with com pletely divided nasals and more than 3 5 5 scales in the mid-dorsal series, the Costa Rican specimen appears to represent an undescribed population to be known a!;
Thorax: uniformly dark brown. Legs dark brown; fore, mid knees, broad base and apex of hind tibia pale; hind tibial comb with six spines, second from spur lon- gest. Wing (Fig. 3) length 1.23-1.58 (1.39, n = 3) mm; width 0.55-0.65 (0.60, n = 3) mm; CR 0.66-0.69 (0.68, n = 3); with contrasting pattern; crossvein r-m faintly darkened anteriorly; most of second radial cell in pale spot; R 3 faintly darkened up to the point where it turns abruptly forward to meet costa; distal pale spot in r 3 transverse, reniform, barely abutting wing margin; M 2 straddled by pale spot nearly its midlength; two distal
5 to 6. Canthals are elongated and slightly pig- mented. One individual (UCR 3359) lacks a postcanthal, the others possess one. Intercanthal count differs among individuals, ranging from 6 to 9 scales. In all examined specimens, internasals are not separated total- ly, although one scale is usually inserted between them. Ventral scales are not different between males and females examined, as 138 and 144 were observed for both sexes. On the other hand, undivided subcaudals in males are 30 and 33, whereas in both females 26 sub- caudals were counted. Interrictals range from 25 to 29. Dorsal scales rows are consistently higher in the neck and midbody than at the vent: observed formulas are 25-25-20, 25-24- 19, 27-24-19, 25-25-19. Nasofrontals range from 23 to 29 scales. One or two oculabials were observed. The number of dorsal blotch- es in the body is 15 and 16, and in all speci- mens more than 70% of the blotches are not fragmented.
Diagnosis: A small, yellowish-tan, short legged member of the Norops fuscoauratus species group characterized by having enlarged posteloacals and a large, bicolored pink and orange dewlap in males and a small, plain dewlap not contrasting in color with the rest of the throat in females.
Thus far, 26 species are known in the genus Cyllopsis Felder; and its geographic distribution is from the Southern United States to Panama. The majority of the species occurs between Guatemala and Mexico. The habitats of these butterflies are described as montane or submon- tane; a few are found in the lowlands (Miller 1974). In CostaRica, five speciesof Cyllopsis have been reported, three of which are known to be endemic to the Talamancan ranges. The host
The shallow-water cumaceans of the west coast of the Americas are not well known (Jones 1969); specifically, only six species have been described from the Pacific coast of the Ameri cas between California and Tierra del Fuego. The animal described herein was taken during a survey of the tidal flat fauna near Punta Mo rales on the Golfo de Nicoya, CostaRica (Var gas 1987), where it shows a seasonal abundance pattern (Vargas 1989). As will be shown below, this newspecies is especially significant as it has sorne of the diagnostic features of the both Family Leuconidae and the Subfamily Bodotriinae of the Family Bodotriidae.
Abstrad: Twospeciesof Clerini (Cleridae: Clerinae), Colyphus hansoni and Enoclerus (E.) puravida , are described from malaise trap samples collected in a patch of c1oudforest at Zurquí de Moravia, San Jose Province, CostaRica. Colyphus hansoni is compared to its congeners and to the sympatric Enoclerus (Coniferoclerus) subviolaceus (Gorham). The newspecies is distinguishable from its congeners on the basis of color, seta! pattem and elytral sculpturing. A sympatric, undescribed Colyphus species resembles C. hansoni in details of shape and sculpturing, and may prove to be its sister species. Colyphus hansoni has the elyra tricolorous (red, ivory and black) and thus is easily separated from the undescribed species which has the e1ytra strictly bicolorous (stramineous and black). The generic status of the latter is discussed in relation to Colyphus. The presence of sexually dimorphic tarsal claws in Colyphus is noted for the first time. Enoclerus (E.) puravida is characterized as part of a complex of several similar and possibly related species distributed in Panarna andCostaRica. It is similar lo several other Mexican and Central American Enoclerus species lhat share small size, ant-like form, shining black or reddish elytral integument and distinctive sculpturing of the elytral base. This group, consisting of E. (E.) tubercularis (Gorharn 1882), E. (E.) gibbus Ekis 1976, E. (E.) albosignatus Ekis 1976, E. (E.) puravida and sorne undescribed species, may eventually prove io form a clade-and thus warrant elevation to subgeneric or generic status. Among described species, E. (E.) puravida is most similar to the PananlalÚan E. (E.) albosignatus, from which it differs by having the of each elytron coarsely alveoJate-¡iunctate and only incidentally and feebly costate, rather than smooth and "embossed" with !bree shalJow carinae. The twospecies also differ in details of coloration and pronotal sculpturing. E. (E.) puravida may be mimicking ants.
Abstract: A new marine gastropod speciesof the genus Trapania Pruvot-Fol, 1931, is described from Cabo Blanco, Puntarenas, Ricaandfrom Islas Secas, Panamá. Trapania inbiolica sp. nov. has a white body with red patches, white rhinophores with sorne liule red patches, yellow appendages with partially red bases. The radula is composed of 28 rows of teeth. Each tooth has a large conical cusp with 21-24 denticlcs. Two or three of those denticles on the inner side of the eusp are smaller than the others. There are also 1-3 small denticles on
compressed antero-posteriorly. Medial teeth of outer row compressed in young, but nearly round in cross section, and blunted, in large adults. All jaw teeth with brown tips in adults. Each upper pharyngeal bone with convex surface bearing oval patch of curved retrorse teeth; anterior teeth largest and reund in cross section, posterior teeth progressively smaller and late rally campressed. Lower pharyngeal plate triangular with apex directed anteriorly and tooth-bearing surface concave; teeth antrorse, small anteriorly and large posteriorly. M.any smaller pharyngeal teeth with shoulder or incipient second cusp on inside of curved surface. Thus, the pharyngeal miU is an efficient shredding apparatus with large teeth opposing small teeth and the upper teeth directed posteriorly and the lower teeth anteriorly.
STAGE 25.- Tadpoles having some yolk present, although gut beginning to form; external gills lost; sinistral, ventro-lateral spiracle present; external nares present. At mid-length of tail depth of caudal musculature les s than one-third depth of tail; caudal musculature distinctly curved upwards posteriorly (Fig. 1 1 ) . Top of head, orbital region, and abdomen moderately pigmented; caudal muscu lature and fins, except anterior one-third of ventral fin, sparsely pigmentedá small flecks present on caudal fin. Mouth, except median anterior edge, bordered by two rows of small papilla e lateraUy and one row posteriorly; both beaks moderate ly developed and bearing small serrations; two upper and three lower tooth rows; second upper row broadly interrupted medially; first lower row slightly shorter than upper rows; second lower row as long as first and interrupted medially: third lower row short and composed of small teeth (Fig. 1 5 ) .
DESCRIPTION : Considerably more variation can be observed between Pacific slope populations of the species than between Atlantic forms. Even within the same river system, the Río Grande de Térraba, the San Isidro del General population varies remarkably frorn Golfito specimens (Fig. 9 ) . This variation consists of differences in body depth, least depth of caudal pedunde, lengt!1 of dorsal and anal fin bases and subtle color pattern differences. Other body proportions and meristic characters are constant between the two populations. If Regan had based C. lethrinus on specimens from San Isidro del General and no other specimens were available except Meek' s Turrialba collection, one would not hesitate in recognizing the two distinct species. Table 4 sums uI' several characteristics which are shared by all Pacific populations and all Atlan tic populations and reveals how the highland material from the Valle Central is intermediate to both of these. The Valle Central specimens, although in habiting the headwaters of one of the large Pacifie drainages, exhibit character istics which are otherwise onl5' observed on Atlantic examples (3 and 4, Table 4) . Regardless of whether these similarities reflect genetie flow in the present or recent past between Atlantic and Pacifie slope populations at this point or are the result of convergence of characters due to sorne environmental factors, 1
Abstract: A newspeciesof fern of the genus Pteris (Filicales: Pteridaceae) endemic to CostaRica. The new fern species Pteris herrerae A. Rojas & M. Palacios, endemic to CostaRica, is described. It differs from P. decurrens C. Presl in basal segments reduced to 1/5-1/2 of the next segment (vs. 2/3-3/4), basal pinnae not bifurcated (vs. bifurcated), pinnae apex mucronate (vs. acuminate) and segment apex undulate (vs. dentate). It differs from Pteris consanguinea in the elliptic pinnae (vs. oblong), two segments reduced on the base (vs. lack), segments entire to undulate (vs. dentate), basal pinnae without basiscopic lobes (vs. with basiscopic lobes) and segment apex entire to undulate (vs. dentate). Rev. Biol. Trop. 54 (3): 1061-1066. Epub 2006 Sept. 29. Key words: Pteris, newspecies, Pteridaceae, ferns, CostaRica.
triton and study of previously unavailable ra diographs of the holotype of diminuta lead us to reassign the species to Bolitoglossa. The rela tively stout and expanded tenninal phalanges are like those of Bolitoglossa rather than resem bling the slender and more pointed elements typical of Nototriton. Furthennore , diminuta lacks a sublingual fold, the absence of which is a disgnostic feature of Bolitoglossa among the neotropical genera. The caudosacral region is typical of Bolitoglossa alpha (Wake and Lynch, 1 976) , with elongate and anteriorly swept transverse processes on the first caudal vertebra. Finally, the trunk vertebrae do not have the relatively compact, nearly opisthocoelous cen tra which are usually present in Nototriton.
subtruncate; nostril small, labial protuberances of nasolabial groove very small; canthus rostralis indistinct, gently rounded. Standard length 6.6 times head width; standard length 4.6 times snout-gular fold length. Deep, distinct gro ove below eye extends full length of eye opening, does not communicate with lip. Eye of moderate size, not greatIy protuberant. Poorly defined postorbital groove extends posteriorly from eye as shallow, realtively broad depression for 1 . 7 mm, proceeds sharply ventrally from posterior terminus and extends across guIar area as well-marked groove parallel to and 3.5 mm anterior to guIar fold. Vomerine teeth 1 1 - 8, in curved series that extend to center of internal nares. Maxillary teeth 2 1 - 22, extending posteriorly to point equal to three-fourths distance through eyeball. Four premaxillary teeth, none piercing lip. Body relatively slender, slightly desiccated by preservation. Very slender tail 1 . 1 times standard length; lateral compression moderate; slight basal constriction. Post-iliac glands small, poorIy defined. Limbs moderately long, slender; when appressed to sides of trunk, two costal folds remain uncovered (limb interval 2 ) ; standard length 4.4 times right fore limb; standard length 3.9 times right hind limbo Digits of hands and feet easily distinguishable but extensively webbed; digital tips broad ly rounded; borders of adjacent digits slope proximally and meet at level of articulation of penultimate and terminal phalanges. Area of webbed pad rela tively large in relation to size of hands and feet, but digital tips remain free from web. Hands and feet flattened; digits flattened, not cylindrical. Subter minal pads distinct, moderate in size; partiallY obscured by flattening during preservation. F ingers in order of decreasing length: 3, 4, 2, 1 ; toes in order of decreasing length : 3, 4, 2, 5, 1 .