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Marc S. Appelhans was born on the 7th of July 1980 in Winterberg, Germany. He attended
high-school in Winterberg and received his Abitur (A-levels) in 2000. After high-school, he
accomplished his civil service at the Sankt Franziskus Hospital in Winterberg.
Marc started his studies in Biology at the Philipps-University in Marburg in September 2001
and started to specialise in Systematic Botany after the Vordiplom (intermediate examina-
tion) in 2003. Next to Systematic Botany as a major, he chose Molecular Cell Biology, Plant Physiology and Physical Geography as minors. During his studies in Marburg, Marc gained
his first experience in teaching. He was recruited as a student assistant for the ‘plant identi-
fication course’ led by Prof. Weber in 2004 and 2005. Furthermore, he frequently led guided
tours in the Botanical Garden of Marburg. During his time in Marburg, Marc took part in
a series of botanical excursions in Germany, and broadened his knowledge on fieldtrips to
the Alps (Austria), the Mediterranean (Mallorca, Malta), South America (Argentina, Chile) and Namibia. For his diploma thesis (equivalent to MSc), Marc embarked on a study about the mycorrhizal structures in the rue family (Rutaceae), supervised by Prof. Weber and Dr. Imhof. He graduated in April 2007 and published parts of his diploma thesis in the journal
Mycorrhiza. From July until October 2007, Marc worked as a scientific assistant in the group of Prof. Weber, working on endemic species of the Maltese Islands.
Marc’s fascination for the Rutaceae family brought him to the Netherlands, where he started his PhD at the Hortus botanicus Leiden/ NHN Leiden in February 2008 under the supervi- sion of Prof. Erik Smets and Dr. Paul Keßler. In the four years of his PhD studies, he worked on several anatomical structures, molecular phylogenetic analyses and the historical bio- geography of the Rutaceae subfamily Spathelioideae. During his time in the Netherlands, Marc gained further teaching experience (1st year students courses; Course on South-East Asian plant families; Course on tropical plant families), led guided tours and held public lectures in the Hortus botanicus and visited Herbaria in London (K), Berlin (B), Jena (JE), Hamburg (HBG), Copenhagen (C), Singapore (SING), Kota Kinabalu (SNP), Havana (HAC),
and Kingston (UCWI, IJ). Marc conducted fieldwork in Jamaica and Malaysia (Borneo), and
he presented his research at international conferences in Leiden, Singapore, Berlin and Mel- bourne. He received prizes for the best student presentations at the PhD day of the Dutch ´research school biodiversity´ 2010 in Leiden, and the international ´Biosystematics´ confer- ence in Berlin in 2011.
In February 2012, Marc will start a one year postdoctoral position at the Smithsonian Institu- tion in Washington DC, USA, extending his studies on the phylogeny and biogeography of Rutaceae.
Van der Weide JC, Ronse De Craene LP, Appelhans MS, Smets E. Floral ontogenetic studies
of selected Rutaceae species with special emphasis on the nectaries. In preparation for Journal
ofSystematics and Evolution.
Stock NK, Boschetti N, Herzog C, Appelhans MS, Niedrig M. The Phylogeny of Yellow Fever
Virus 17D Vaccines. Submitted to Vaccine.
Appelhans MS, Keßler PJA, Smets E, Razafimandimbison SG, Janssens SB. Age and historical biogeography of the pantropically distributed Spathelioideae (Rutaceae, Sapindales). Resub-
mitted after revision to Journal of Biogeography.
Appelhans MS, Smets E, Razafimandimbison SG, Haevermans T, van Marle EJ, Couloux A, Rabarison H, Randrianarivelojosia M, Keßler PJA. 2011. Phylogeny, evolutionary trends, and
classification of the Spathelia / Ptaeroxylon clade: morphological and molecular insights. An-
nals of Botany 107: 1259-1277.
Razafimandimbison SG, Appelhans MS, , Rabarison H, Haevermans T, Rakotondrafara A,
Rakotonandrasana SR, Ratsimbason M, Labat J-N, Keßler PJA, Smets E, Cruaud C, Cou- loux A, Randrianarivelojosia M. 2010. Implications of a molecular phylogenetic study of the
Malagasy genus Cedrelopsis and its relatives (Ptaeroxylaceae). Molecular Phylogenetics and
Evolution57: 258-265.
Appelhans MS, Smets E, Baas P, Keßler PJA. 2010. Cneorum (Rutaceae) in Cuba? The solu-
tion to a 150 year old mystery. Taxon59: 1126-1134.
Appelhans M, Weber HC, Imhof S. 2008. Rutaceae sampled from Germany, Malta, and Mal-
lorca (Spain) are associated with AMF clustering with Glomus hoi Berch & Trappe. Mycor-
It all started with a missed train. I was standing at a platform at the train station in Cologne and knew that I would be late for my job interview in the botanic garden of Leiden Univer-
sity. I thought I had lost the job before even getting it, but it turned out to be OK and I finally
started my PhD project in February 2008. The first people I have to thank are therefore Erik
Smets, Paul Keßler, Johan Mols and Marco Roos, the members of the committee that decided to hire me despite this bad start.
This thesis would be much less satisfying without the input of Sylvain Razafimandimbison
and Steven Janssens, who were of great help in theoretical and practical questions concerning phylogeny and biogeography. I´m glad and honoured that my name appears on publications together with yours.
Help and support came from several directions. I would like to thank Bertie Joan van Heuven, Marcel Eurlings, Natasha Schildo, René Glas, Erik Jan van Marle and Edith Kapinos for their help the anatomical and molecular labs; Max van Balgooy, Willem and Brigitta de Wilde, Wil-
lem Prud´homme van Reine, Phongsak Phonsena, Hajo Esser, Jackie Kallunki, Wayt Thomas,
Ramona Oviedo, Pável Oriol Rodríguez Vásquez and Angel Vale for collecting and sending specimens; Patrick Lewis, Helen Jacobs and George Proctor for their great help during an
unforgettable field trip to Jamaica; Jan Frits Veldkamp and Werner Greuter for answering my
unexperienced questions about nomenclature; Anita Walsmit Sachs-Jansen for her beauti- ful botanical drawings; Anita and Esmée Winkel for teaching me how to make a botanical drawing myself; Connie Baak for help with InDesign and layout-questions; David Mabberley, Jackie Kallunki, Hajo Esser and Christa Beurton for general discussions and advise concern-
ing my project and Rutaceae; Art Vogel, Rogier van Vugt and the staff of the Hortus botanicus
for taking care of my plants in the greenhouses; Lisa Becking, Liberty Blank, Atiyo Ghosh and Nicole Webster for correcting and proofreading the English of my manuscripts.
Brigitta and Willem de Wilde are thanked for ´de heel gezellige en leuke tochtjes´. Without you, I would surely have felt less at home in Leiden and I would have seen less from the Nether lands.
Many thanks go to my fellow students and friends from NHN, naturalis, CML, DNAmarker- point, LEAD as well as visiting students. Especially Kanchana Pruesapan and Christian Böde- ker are thanked for help in the lab and many discussions about my project.
Life consists of more than only botany. Thanks to all my friends from Marburg, Leiden and all
over the world for showing me that.
Großer Dank gebührt natürlich meiner Familie für die Unterstützung während meiner Stu- dienzeit in Marburg und Leiden.