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II. Consideraciones de la perspectiva comparada

3.3. Derecho a la unión de hecho

Gallardoneris iberica Martins, Carrera-Parra, Quintino & Rodrigues, 2012 Notes: Reported from Greece by Katsiaras et al. (2017). In the Mediterranean also known from the central and western basins (Bertasi et al. 2014, Gómez et al. 2015, Mikac 2015, D’Alessandro et al. 2016). Its presence in the region was probably overlooked until its description.

Hilbigneris gracilis (Ehlers, 1868)

Nomenclature: Hilbigneris gracilis (Ehlers, 1868) | Lumbriconereis gracilis Ehlers, 1868 | Lumbrineris gracilis (Ehlers, 1868)

Notes: Type locality: Mediterranean (Adriatic). Lumbricalus adriatica (Fauvel, 1940)

Nomenclature: Lumbricalus adriatica (Fauvel, 1940) | Lumbrineris adriatica (Fauvel, 1940)

Notes: Type locality: Mediterranean (Adriatic). Carrera-Parra (2004) redefined the genus, including Lumbricalus adriatica, using the maxillary apparatus in combination with chaetal types for differentiating species.

Lumbrinerides acuta (Verrill, 1875) sensu Ramos, 1976

Nomenclature: Lumbrineris acuta Verrill, 1875 | Lumbrinerides acuta (Verrill, 1875) Notes: Questionable status. Reported from Greece by Dounas (1986). The record refers to Lumbrinerides acuta sensu Ramos 1976. The original description by Verrill

(1875) is not detailed, but the redescription by Perkins (1979) based on specimens collected in the type locality (Rhode Island, USA) includes an accessory tooth on maxillary pair MI. The absence of an accessory tooth, as it is described in Ramos (1976), may indicate an undescribed species (Gómez et al. 2015). In the Mediterranean also known from Spain (Sardá et al. 1999), Italy (Castelli et al. 2008), Turkey (Çınar et al. 2014) and the Adriatic (Mikac 2015), but it is unclear whether these records refer to specimens having an accessory tooth or not. While it is possible that the species occurs in Greece, it has so far only been reported in grey literature and has not been found in over three decades, therefore its presence in Greece is here considered questionable.

Lumbrinerides amoureuxi Miura, 1980

Notes: In the Mediterranean known from Greece, Turkey (Çınar et al. 2014) and Cyprus (Çınar 2005), otherwise distributed along the Atlantic coast of Europe.

Lumbrinerides carpinei (Ramos, 1976)

Notes: New record for Greece. Two specimens, Pachia Ammos, Crete, 35°06'39.6"N, 25°48'32.4"E, 1–5 m depth, fine to coarse sand, collected in the framework of the MEDCORE project (unpublished data from Papageorgiou et al. 2006). Literature used for identification: Ramos (1976). Type locality: Western Mediterranean.

Lumbrinerides laubieri Miura, 1980

Notes: In the Mediterranean only reported from Greece (Simboura and Zenetos 2005). Known from deep waters off the Atlantic coast of Europe.

Lumbrinerides neogesae Miura, 1981 Native status: Non-native (casual)

Notes: New record for Greece. One specimen, Elafonisi, Crete, 35°16'20.7"N, 23° 32'15.9"E, 1 m depth, fine sand, collected in the framework of the MEDCORE project (unpublished data from Papageorgiou et al. 2006). Literature used for identification: Miura (1980), Gravina and Cantone (1991). In the Mediterranean also known from Italy (Gravina and Cantone 1991). Originally described from South Africa.

Lumbrineriopsis paradoxa (Saint-Joseph, 1888)

Nomenclature: Lumbriconereis paradoxa Saint-Joseph, 1888 | Lumbrineriopsis paradoxa (Saint-Joseph, 1888) | Lumbrineris paradoxa Saint-Joseph, 1888

Lumbrineris coccinea (Renier, 1804)

Nomenclature: Lumbriconereis coccinea (Renier, 1804)| Lumbrineris coccinea (Renier, 1804)

Notes: Type locality: Mediterranean. Carrera-Parra (2006) considers records from outside the Mediterranean Sea doubtful.

Lumbrineris inflata Moore, 1911

Notes: Questionable status. Reported from Greece by Arvanitidis (2000a) and Kitsos (2003); the first Mediterranean record was by Giangrande et al. (1981). Arvanitidis (2000a) argues that the species may have been underreported in the Mediterranean due to its similarity with Lumbrineris coccinea (Renier, 1804). Carrera-Parra (2006) and D’Alessandro et al. (2016) consider Mediterranean records of the species doubtful based on its otherwise East Pacific distribution. Çınar (2009) argues that the description and illustrations of Lumbrineris inflata by Giangrande et al. (1981) fit the characters of Lumbrineris perkinsi Carrera-Parra, 2001. Lumbrineris inflata differs from Lumbrineris perkinsi mainly in the number of teeth of maxilla III (M III). Giangrande et al. (1981), as well as Arvanitidis (2000a) for Greece, report a maxillary formula which could correspond to Lumbrineris perkinsi ("M III with 3–4 teeth"). Although published descriptions and records from the Mediterranean indicate the presence of Lumbrineris perkinsi, re-examination of all the material is needed to draw conclusions about the identity of the Greek records.

Lumbrineris latreilli Audouin & Milne Edwards, 1834

Nomenclature: Lumbriconereis latreilli Audouin & Milne Edwards, 1834 | Lumbrineris latreilli Audouin & Milne Edwards, 1834

Notes: Frequently reported from Greece, however, mainly on the basis of external and unreliable diagnostic characters. Species descriptions focusing on the maxillary apparatus and other unambiguous and size-independent characters have recently become available (Carrera-Parra 2006, Martins et al. 2012, D’Alessandro et al. 2016). Therefore, re-examination of material could reduce the frequency of records and even reveal the presence of overlooked species. In the Mediterranean, the species is widely recorded (e.g. Campoy 1982, Çınar 2005, Castelli et al. 2008, Ayari et al. 2009, Çınar et al. 2014, Mikac 2015). Otherwise distributed in the North-East Atlantic; its presumed cosmopolitan distribution is questionable (Carrera-Parra 2006).

Lumbrineris luciliae Martins, Carrera-Parra, Quintino & Rodrigues, 2012

Notes: New record for Greece. Specimens previously reported as as Lumbrineris cingulata Ehlers, 1897 by Chatzigeorgiou et al. (2016) from rocky shores in northern Crete were re-examined and found to belong to Lumbrineris luciliae as described by

Martins et al. (2012), although the reported slight distal curvature of the aciculae could not be observed. In the Mediterranean also known from Italy (D’Alessandro et al. 2016). Otherwise known from the Atlantic coast of the Iberian Peninsula (Martins et al. 2012). Lumbrineris nonatoi Ramos, 1976

Notes: Type locality: Mediterranean (Baie de Rosas, Spain). Lumbrineris nonatoi has several diagnostic characters in common with the – later described – Gallardoneris iberica Martins, Carrera-Parra, Quintino & Rodrigues, 2012. Due to this confusion, Bertasi et al. (2014) and Katsiaras et al. (2017) suggest that several literature records of Lumbrineris nonatoi could belong to the overlooked Gallardoneris iberica, although the overall presence of the former in the region has not been doubted.

Ninoe armoricana Glémarec, 1968 Ninoe nigripes Verrill, 1873

Nomenclature: Ninoe kinbergi Ehlers, 1887 | Ninoe nigripes Verrill, 1873

Notes: Questionable status. Reported from Greece as Ninoe kinbergi by Vamvakas (1970) and Vamvakas (1971). Arvanitidis (2000a) lists these records under the name Ninoe nigripes, following the synonymy established by Pettibone (1963). In the Mediterranean reported from the central basin as Ninoe kinbergi (Castelli et al. 2008) and as Ninoe nigripes by Mikac (2015) from the Adriatic Sea. Otherwise distributed in the West Atlantic.

Scoletoma emandibulata (Pillai, 1961)

Nomenclature: Lumbrineris emandibulata Pillai, 1961

Notes: Questionable status. Reported from Greece by Dando et al. (1995) and Akoumianaki (2004). The record by Akoumianaki (2004) actually refers to Scoletoma emandibulata mabiti, as Ramos 1976 is given as the authority; it remains unclear whether the same holds true for the specimen of Dando et al. (1995). Originally described from the Indian Ocean.

Scoletoma emandibulata mabiti (Ramos, 1976)

Nomenclature: Lumbrineris emandibulata mabiti Ramos, 1976 Notes: Type locality: Mediterranean (Catalan Coast).

Scoletoma fragilis (O.F. Müller, 1776)

Nomenclature: Lumbriconereis fragilis (O.F. Müller, 1776) | Lumbrineris fragilis (O.F. Müller, 1776) | Scoletoma fragilis (O.F. Müller, 1776)

Notes: Species complex. Frequently reported from Greece and other parts of the Mediterranean (Castelli et al. 2008, Ayari et al. 2009, Çınar et al. 2014, Mikac 2015). Otherwise distributed in the Atlantic Ocean, Arctic Ocean and North Sea. However, molecular analyses revealed the existence of at least two putative cryptic species, one in the Atlantic and one in the Arctic Ocean (Carr et al. 2011).

Scoletoma funchalensis (Kinberg, 1865)

Nomenclature: Lumbriconereis funchalensis Kinberg, 1865 | Lumbrineris funchalensis (Kinberg, 1865) | Scoletoma funchalensis (Kinberg, 1865)

Notes: In the Mediterranean also known from the western and central basins (Campoy 1982, Castelli et al. 2008), Tunisia (Ayari et al. 2009), Turkey (Çınar et al. 2014) and the Adriatic (Mikac 2015). Otherwise distributed in the North-East Atlantic. However, Oug (2011) noted that Scoletoma funchalensis is poorly known and the described characters by Fauvel (1923) could be confused with juveniles of several species. Scoletoma impatiens (Claparède, 1868)

Nomenclature: Lumbriconereis impatiens(Claparède, 1868) | Lumbrineris impatiens Claparède, 1868 | Scoletoma impatiens (Claparède, 1868)

Notes: Type locality: Mediterranean (Gulf of Naples). George and Hartmann-Schröder (1985) synonymised Scoletoma impatiens with the South African Scoletoma tetraura (Schmarda, 1861), but several authors do not accept this synonymy (see also notes under Scoletoma tetraura).

Scoletoma rovignensis (Fauvel, 1940)

Nomenclature: Lumbrineris rovignensis Fauvel, 1940 | Scoletoma rovignensis (Fauvel, 1940)

Notes: Questionable status. Reported from Greece by Dounas (1986). Type locality: Mediterranean (Adriatic). While it is possible that the species occurs in Greece, it has so far only been reported in grey literature and has not been found in over three decades, therefore its presence in Greece is here considered questionable.

Scoletoma tetraura (Schmarda, 1861)

Nomenclature: Lumbrineris tetraura (Schmarda, 1861) | Scoletoma tetraura (Schmarda, 1861)

Notes: Questionable status. Reported from Greece by Koukouras and Rousso (1991). In the Mediterranean also known from Turkey (Doğan et al. 2005; but considered questionable by Çınar et al. 2014), Cyprus (Çınar 2005), Italy (Castelli et al. 2008), Albania (Marzano et al. 2010) and the Adriatic (listed under Scoletoma impatiens (Claparède, 1968) by Mikac (2015)). Originally described from South Africa. George and Hartmann-Schröder (1985) synonymised the European species Scoletoma impatiens with Scoletoma tetraura. However, several authors question the presence of the latter in Europe and suggest using the name Scoletoma impatiens for specimens from the area until a more detailed revision becomes available (Martins et al. 2012, Çınar et al. 2014, Gómez et al. 2015, Mikac 2015, D’Alessandro et al. 2016). The main diagnostic character of the genus can be problematic, since several juvenile specimens of Lumbrineris and Hilbigneris species may lack composite hooded hooks and can be key out as Scoletoma (Oug 2011).

Magelonidae Cunningham & Ramage, 1888