Capítulo II: RESISTENCIA NEGRA EN LA TOMA: MUJERES EN DEFENSA DE
3. Características de la acción colectiva de las mujeres de La Toma
3.2 Motivaciones y repertorios de la acción colectiva
3.2.1 Mujer y defensa de lo propio, una mirada desde la Diáspora Africana
Lectotype m . ( Dr . S . Bleszynski ) in British Museum ( Natural History) , slide BM Pyral 5630, labelled "Mt . Hutt New Zealand /1 /01 R .W . F . Crambus
aethonellus n . sp . "
Holotype m . in British Museum (Natural History ) , slide BM Pyral 5636 , labelled "Mt . Cook New Zealand GVH . /1 2/99 Crambus antimorus11 • Checked against figure by Philpott ( 1 929 , p .495 ) by Dr . S . Ble szynski .
Lectotype m . ( Dr . S . Bleszynski ) In British Museum (Natural Histo�) slide
BM Pyral 5637 , labelled "Mt . Cook New Zealand /1 2/99 GVH . Crambus heteranthe s" . Lectotype m . ( Dr . S . Bleszynski ) in British Museum (Natural Histo�) slide
BM Pyral 7974, labelled " Invercargill New Zealand, A .P . 1 . 07 Crambus saristes" .
Holotype m . in British Museum (Natural Hi story ) , slide BM Pyral 5631 , labelled " Invercargill New Zealand GVH ./00 Crambus saristes" . Checked against genitalia of lectotype aethonellus by Dr . S . Ble szynski .
Lectotype m . ( Dr . S . Ble szynski ) in British Musewn (Natural History ) , slide BM Pyral 7973 , labe lled 11Longwoods New Zealand AP . 2700 ' 1 2 . 1 0 . Crambus meristes" , with 4 paralectotypes .
Probable syntype s of aethonellus m . f . , author' s preps AGP, AGQ in CMNZ labelled
"Mt . Hutt New Zealand Jan 77 Crambus aethonellus n. sp . " and "Mt . Hutt New Zealand Jan 82 Crambus aethonellus n . sp . "
EXTERNAL CHARACTERS ( Slides 2 , 3 ) Detailed de scriptions by Meyrick
( 1 88.3 ; 1 901 ; 1 909 ; 1 91 9 ) . Coloured illustrations by Hudson ( 1 928 , Plate XIX, figs . 7 m . 23 m . Plate XLV, fig . 1 8 f ) and abbreviated de scriptions . Wing span;
1 3-20 mm (m) , 1 5-21 mm ( f ) . Female frenulum triple . This variable specie s lacks the subapical white patch typical of the forewing of the superficially similar o .melitastes . Eye diurnal type ( H � J ... .,.! .. le l <t C6).
MALE GENITALIA ( Fig • 44) : As for generic and subgroup .de scription except : Arms of vinculum about
i
uncus ; saccus tapered with " keel" of Philpott ( 1 929 ) ;costa flanged on inner posterior edge , dorsal margin about � uncus . Sacculus not different iated . Gnatho s tapered , pointed, short , % uncus . Aedoeagus 2t,x uncus , length to median breadth ratio 6-7 : 1 , dorsal midline strong with apical torsion 45° sinistrad , small apical thorn dorso-sinistrad .
FEMALE GENITALIA ( Fig . 1 32 . From female compared with CMNZ probable syntype ) : As for generic de scription except : Eighth abdominal tergite encircling ostium ,
2-3 t ime s as wide in ventral as in dorsal midline . Eighth sternite absent , ductus bursae 4i-5x length of anal papillae , weak. Ductus seminalis joining
1
at 2 . Ostium bursae narrow , dorso-ventrally compre ssed , with exaggerated
dorsal lip , ostiolar pouch not swollen. Corpus bursae with two markedly
unequal oval signa .
LARVA : Not known .
HOST PLANTS : Not known, but possibly red tus sock , Ch . rubra .
DISTRIBUTION ( Fig . 227) : South Island o f New Zealand from sea level in the extreme south , w.hete:: it oc curs among sandhills near Invercargill ( Hudson, 1 928 ) , to 4 , 000 ft . in the mountains of the main axis . Associated with boggy areas ,
adult flight period late October to late January, suggesting one protracted
emergence by a s ingle generation each year.
MATERIAL EX.MHNED : 1 59 m . 23 f .
5 m . no· date or locality, CM , R.W . Fereday .
Beaumont : 2 m . NBH, 2 f . NBC , �' 7 . 1 2 . 63 , FRI , J . S . Dugdale . Cass , Cant . : 3 m . Nov . 1 924, CM, A . Tonnoir .
Danssy ' s Pas s , 3 , 067 ft . : 1 m . 2 f . NBI , NBJ , 1 . 1 2 . 63 , FRI , J.S . Dugdale . Drayton Station, Cant . : 1 m . 6 . 1 1 . 1 879 , CM, R.W. Fereday .
Flagstaff , Dunedin: 5 m . 1 f . 1 2 . 1 . 1 5 , ED , A . Philpott .
Gore Hill : 3 m . MAD , �' MAG , 1 f . �' 9 . 1 .45 , DM, J.T . Salmon .
Jack' s Pass, Hanmer : 6 m . 1 5-22 . 1 1 .32 , CM , S . Lindsay.
Lake Ohau: 4 m . 26 . 1 2 . 35 , CM, S . Lindsay .
Lake Tekapo : 1 0 m . 1 f . Eb , 4 m . CM , 24. 1 2 . 28 ; S . Lindsay .
Leith : 2 m . 27 . 1 1 . 21 , AM, C .E . Clarke .
Longwood s , Sthld . : 1 m . 1 f • .!:M, DM, 6 m . ED , 5 . 1 2 . 1 3 , 2 m . M.AA , RAB , DM ,
4 m . ED , 20. 1 2 . 1 5 , 1 m . �' 1 f . �' 1 . 1 . 1 6 , DM; A . Philpott .
Maruia Springs : 2 m . 22 . 1 2 . 39 , CM, S . Lindsay .
Mt . Cook, 3 , 000 ft . : 3 m . Q£Q, GFH, 1 5-1 8 . 1 2 . 1 899 , DM, G .V . Hudson;
6 m. ED , 4 m. CM , 3-4 . 1 . 29 , S . Lindsay; 3 m. SAA , SAC , SAD , 2 f . SAB , GFM , DM , 9 m . ED , 1 1 m . 1 f . CM, 1 2 . 1 2 . 29 , A. Philpott .
Mt . Grey , Cant . : 3 m . 1 f . CM , 6 m . ED , 1 1 . 1 1 .23 , S . Lindsay; 1 m . CM,
1 f . ED , 1 4-1 5 . 1 1 . 25 , S . Lindsay; 1 m . 1 f , ED , 2 m . CM, 30 . 1 0 . 27, S . Lindsay; 1 m . 1 f , 8 . 1 . 29 , CM , 2 m . 3 . 1 1 . 29 , CM , S . Lindsay .
Mt . Hutt , Cant . : 1 m . AGP, Jan. 1 877 , CM , 1 f , Jan . 1 882 , CM, R .W . Fereday. Mt .Mangatua , 2 , 1 00 ft . : 1 m . �' 1 0 . 1 2 . 63 , FRI , J . S . Dugdale .
Mt . Peel : 3 m . 27 . 1 . 2 7 , CM, S . Lindsay .
Mt . Wakefield , 4-5 , 000 ft . : 1 f . JBA , 1 4 . 1 . Q� , FRI , J . S . Dugdale . Nevis : 4 m . 5 . 1 1 . 1 0 , AM , C .E . Clarke .
Omao hut : 1 f . 7 . 1 2 . 63 , FRI , J . S . Dugdale .
Peel Fore st : 2 m. Nov . 1 920, ED , A . Philpott ; 6 m . 20 . 1 1 . 30 , CM, S . Lindsay . Puhi-puhi , Marl . : 1 m . 1 f . 3-6 . 1 2 . 59 , ED , E .S . Gourlay .
Seaward Mo ss , Sthld . : 1 m . Bt;f, 1 f . RAA , DM, 2 m. ED , no date , A . Philpott ; 2 m . 2 7 . 1 . 1 5 , ED , A . Philpott .
Silverstream, Otago : 1 m . �' 1 4 . 1 2 . 1 5 , DM, 7 m . AM , C . E . Clarke . Table Hill , Sthld . : 1 m . 30 . 1 2 . 1 3 , ED , A . Philpott
Takitimu Mountains : 2 m . 28 . 1 2 .1 7 , ED , A . Philpott .
Waitaki , 2 , 000 ft . : 1 f . �' 1 .12. 63, FRI , J . S . Dugdale .
Waitati : 1 m . 7 . 1 2 . 1 6 , DM, G .V . Hudson; 4 m . 1 6 . 1 1 . 1 7 , AM, C .E . Clarke We�erburn : 1 m . no date , ED , G . Howes .
White Rock, Cant . : 1 m . Nov . 1 91 9 , CM , 1 f . ft£, 26 . 1 1 . 22 , DM, S . Lindsay .
REMARKS : This variable specie s was de scribed six time s by Meyrick ( 1 882 - 1 91 9 ) .
The form sariste s has a median constriction in the transverse white fascia of the forewings , but this is neither a constant or reliable character for the type locality . Hudson ( 1 92S) expressed doubts concerning the specific status
of saristes . Similar doubts concerning the status of auliste s and meritus
made by ?hilpott ( 1 9>29 ) . There i s no geographical separation which could be used to justify subspecific status . Form sariste s , form meristes and form aulistes have been taken near Invercargill at the same locality as
typical aethonellus , although it must be noted that Meyrick ( 1 91 9 ) insisted with
unpublished remarks by A . Philpott that they were all the same specie s .
In the Hudson collection in the Dominion Museum , Wellington the specimens labelled heteranthe s are larger than those labelled saristes , but such a size difference between alpine and lowland specimens is conunonly found in other members of this genus , for example and flexuo sellus . antimorus is characterised by white hindwing s and a paler brown forewing
Form
colour than the other forms , but although this form appears to be c onfined to the Mount Cook· and Lake Tekapo areas , a series of specimens taken by
S . Lindsay and A . Philpott at both localitie s include all graduations in
hindwing colour from pure white t o the dark glossy brown o f typical aethonellus . Dr. S . Ble szynski has informed me
(
in litt .)
that the genitalia of syntypemales of each form in the Briti sh Museum are ident ical . Differenc e s in the
genitalia of male saristes , meristes , antimorus and heteranthes shown by Philpott ( 1 929 ) , are very slight and fall well within the range of variation found by me for the specie s by examining far more material than was available
to Philpott , or than he was prepared to examine . There are some point s of inaccuracy in his drawings , e specially in the shape of the vinculum and saccus . However they are minor and due to the preparations being compressed at slightly different angle s on the slide s and be ing drawn apparently without compensation
fo r this .
Although the foodplant of this species is unknown, the known di stribution of aethonellus is coincident with that shown for red tussock
Chionochloa rubra
(
Burrows , 1 967) . His remarks about the sites of rubra inthe alpine zone also sugge st a relationship ; 11 • • are very poorly-drained , flat
to gently-sloping areas on cirque floors or flat-topped ridges up to about • • • in wetter sites Ch .rubra vegetation grades into bog . "
Orocrambus catacaustus
(
Meyrick)
Crambus catacaustus Meyrick , 1 885 , Trans . N . Z . Inst . , 1 7 , p . 1 34 . Orocrambus catacaustus Meyrick, 1 888 , Trans . N . Z . Inst . , 2 0 , p . 67 .
Orocrambus Meyrick, 1 91 2 , Trans . N.Z . Inst . , 44, p . 1 1 8 .
(
Syn .Ble szynski & Collins , 1 962
)
I choose as lectotype a syntype··m in CMNZ , author ' s prep . �GJ , labelled 11Arthur ' s
Pass New Zealand 3 , 0001 25
/
1/
83 RWF, Crambus catacaustus" 4 other syntype s inBritish Museum
(
Natural History)
have the same label data .Lectotype m
(
Dr . S . Bleszynski)
in Briti sh Museum(
Natural History)
and 1 parale ctotype labelled 11L . Wakatipu New Zealand GVH. 2 . 1 1 . Orocrambus Checked against catacaustus by Dr . S . Ble szynski .EXTERNAL CHARACTERS
(
Slide4) :
Detailed descriptions by Meyrick(
1 888;1 91 2
)
. Coloured illustrations by Hudson(
1 928 , Plate XX, figs . 24 m , 25 m)
,and abbreviated descriptions. Wing span 21 -2� mm
(
both sexe s)
. A largerinsect than but be st separated with certainty by an examination of genitalia . Female frenulum triple , but with two elements occasionally
partly fused for about basal t. Eye diurnal type .
MALE GENITALIA
(
Fig .45 ) :
As for generic and subgroup de scription except :Arms of vinculum about i-� uncus ; saccus apically truncate � slightly
excavate ; sacculus with small flange ; gnathos tapered, point ed ; aedoeagus
stout , 2
!-
2�
uncus , l ength to median breadth ratio 6-7 : 1 , do rsal midline s�r6ngwith
45°
sinistrad torsion towards apex , and with a simple thorn in this l ine·1
s from apex .
FEMALE GENITALIA
(
Fig . 1 33 . From topotypic female)
: As for generic descript ionexcept : Eighth abdominal tergite fusing dorsolaterally with ostium bursae , twice as wide at fusion as in.dorsal midline . Anterior apopophyse s repre sented
only by ve stigial pro jections on the anteri or margin of the eighth tergite in the lateral midlirn s . Eighth sternit e repre sented only by two small chitinised
spots in the membrane between the anal papillae and the lamella postvaginalis .
Ductus bursae 3-3
�
length of anal papillae , caudal�
strong , with ventral pleatsbetween � and
�
from o stium . Ostium bursae a simple dorso-ventrally compre ssedfunnel . Corpus bursae with two large oval s igna . LARVA: Not known .
HOST PLANTS : Not known, pos sibly red tussock, Ch.rubra or some alpine bog plant .
DISTRIBUTION ( Fig . 228
)
South Island of New Zealand, associated with bogs .Adult flight period December to early March.
MATERIAL EXAMINED : 91 m . 41+-f . ·
Arthur 1 s Pass , 3 , 000 ft . : 1 m . �� ( lectotype ) , 25 . 1 . 1 883 , CM, R.W . Fereday;
3 .m. CM, 1 .f . ED , ·1 .2.1 5 , G . V. Hudson; 1 m . 7 . 2 . 1 5 , DM, C .E . Fenwick;
1 m. 1 f. 1 7. 1 . 20, ED , A. Philpott ; 1 m. Feb . 1 920, CM, G .V . Hudson;
1 3 m. 2 f . 7�21 . 2� , AM, · C .E . Clarke ; 8 m . 1 f . CM , 1 2 m. 1 f . ED , 6-9 . 2 . 26 ,
S . Linds.�y; 4 m. 23 . 1 . 28 , ED , S . Lindsay; 2 m . �. 2 2 . 2 .40, DM,
J . T . Salmon; 4 m. 1 f .
Mili'
25-28 . 1 2 .40 , CM, S . Lindsay; 1 f. 2 .1 .43 , ED ,E .S . Gourlay .
Billow Mountains : 1 m. 20 . 1 . 20, ED , A . Philpott .
Bold Peak: 2 m . �' 1 f . �' DM, 1 f . ED , 1 1 . 2 . 1 3 , G . Howes ;
1 m . .1 f .
BDC ,
1 0 . 2 .43 , DM, G. Howes .Cascade Creek, Angelus Basin, 5 , 700 ft . : 1 m . 9 . 2 . 64, FRI , J.S . Dugdale .
Gordon' s Pyramid: 1 f . 2 . 3 . 27 , ED , A . Philpott . Homer : 1 m . 28 . 1 2 .41+, DM, G . Howes .
Hump Range : 2 m. 28 . 2 . 1 9 , ED , A . Philpott .
Iron Hill 4,300-5 , 300 ft . : 4 m . 5 f . 8-9 . 2 .66 , FRI , J . S . Dugdale . Keplef. Mountains : 1 m. 3 . 1 . 25 , AM, C .E . Clarke .
Key Ridge , Ailsa, 3-3 ,500 ft . : 9 m . 4 f . 1 1 . 1 . 66 , FRI , J .S . Dugdale .
Key Summit : 1 m . �' 22 . 1 2 .44, DM, J .T . Salmon.
Lake Sylvester Track: 1 f. 1 1 . 2 .66 , FRI , J .S . Dugdale .
Mt . Arthur, 4 , 000-4, 500 ft . : 1 m . 1 f . 1 4 . 1 . 22 , 3 f . 2 . 2 . 23 ; ED , A . Philpott ;
2 m . 1 f . 2-3 . 1 0 .3 6 , ED , G . V . Hudson, 3 m . CM , 1 f . ED , 27-28 . 1 .41+, G . Howes . Mt . Arthur Tableland, 5 , 000 ft . : 1 m . 28 . 1 . 24, CM, S . Lindsay.
Mt . Burns : 1 m. 23 . 1 . 1 4, DM, G . Howe s ; 3 m . 1 f . 1 4·22 ;1.!1 6 , ED , A . Philpott .
Mt . Crystal Ridge , 4, 500-5 ,900 ft . : 2 m . 8 f. 1 4. 2 . 66 , FRI , J.S . Dugdale .
Mt . Gertrude , �rier Stream, 3 , 700 ft . : 1 f . 1 6 . 1 . 63 , FRI , J.S . Dugdale . Mt . Luxrnore , 3 , 500 ft . : 1 m . 5 f. FRI , J . S . Dugdale .
Red Hills : 1 4- . 1212!'. 1 0.2 .64, FRI , J.S . Dugdale .
RE�� : No consistent differences were found in either male or female genitalia of and typical catacaustus forms . Slight variation in
proportional size of structures was noted from specimen to specimen, but
this variation was found in both forms . Meyrick separated from
catacaustus primarily on it s constricted white forewing fascia . Hudson ( 1 951 ) ,
doubted the specific distinctness of and in the long series examined
in this study I found a complete range of intermediate s . J .S . Dugdale
( pers . comm. ) mentions that the form appears to predominate in the
Mt . Arthur region , but there are also typical catacaustus specimens from
this locality in the Canterbury MuseQm collection. Both forms have been taken together at Arthur ' s Pass and the mountains around Lake Wakatipu . I therefore find no grounds at this stage to give even subspecific rank. The constriction of the fascia generally ( although not invariably) occurs in small
dark male s . The syntypic specimens were first examined by Dr. S . Bleszynski
some years ago , and was published as a synonym of catacaustus in the catalogue of world Crambinae by Bleszynski and Collins ( 1 962 ) .
In the phylogenetic scheme set out in this work O . catacaustus is
more closely related to O .aethonellus than to any other species .
Orocrambus dicrenellus (Meyrick) comb . nov.
Crambus dicrenellus Meyrick, 1 882 , New Zealand Journal of Science ( Dunedin) , 1 , P • 1 87 .
Crambus dicrenellus Meyrick, 1 883 , Trans . N.Z . Inst . , 1 5 , p . 22 . Lectotype m and paralectotype m . (Dr. S . Ble szynski ) in British Museum
(Natural History) , slide BM Pyral 7983 . Labelled "Mt . Hutt New Zealand /1/81 RWF; Crambus dicrenellus" .
Probable syntype m. in CMNZ , author ' s preps AGX , AGY , labelled "Mount Hutt , 5 . 1 . 68 RWF Crambus di crenellus n , sp . " and "Drayton Station (Mt . Hutt ) 5 . 1 . 68 RVfl!"' Cram bus dicrenellus n, sp . "
Plate XX , fig . 1 8 " f . " ( although the female i s not known) and abbreviated description. Both the description by Hudson on p . 1 64 and the more detailed description by Meyrick ( 1 883 ) can be assumed to include as well .
Separated at once from the very similar by the conical frons of
the latter . Best separated from by examination of genitalia . Wing span of male 29 - 33 mm .
MALE GENITALIA : ( Fig . 46 compared with probable syntype in CMNZ and with
le ctotype drawing by Dr . S . Ble szynski ) : As in generic and species group
description except for following : Arms of vinculum % uncus , saccus tapered, with " keel" of Philpott ( 1 929 ) . Costa of valve flanged , dorsal margin about
3 7
4 - e uncus . Gnathos tapered, pointed, e qual to uncus . Aedoeagus apically
dilate , 2% - 3x uncus , length to median breadth ratio 1 1 -1 2 : 1 , dorsal midline strong, terminating in a sigmoid spear-shaped apical spur about * total length of aedoeagus .
LARVA : Not known.
HOST PLANTS : Not know�, but possibly Chionochloa australis .
DISTRIBUTION ( Fig . 2 29 ) : Alpine and subalpine tussock grasslands of the central South Island of New Zealand to 4,·ooo ft . Adult flight period
November to February, but .no. December records presently known . Records
given by Hudson ( 1 928 , p . 1 64 include and should be ignored . )
MATERIAL EXAMINED : 1 5 m . / Ball Glacier: Ben Lomond: 1 m . fl!, DM, 2 m . llli:f_, CM, 2 m . ED , 7-1 4.2 .29 , A . Philpott . 1 m. 25 .1 1 . 1 2 , DM, A . Philpott .
Jack' s Pass , Hanmer : 1 m . 8 . 2 .31 , CM, S . Lindsay .
Mt . Cook: 2 m . ABA, BGB, CM, 3 m . ED , 8-1 4.2 . 29 , A . Philpott .
Mt . Hutt , Cant . : 2 m . ill' !f:X, 5 . 1 .1 879 , CM, R .W. Fereday. Sedgmere , 3 ,550 ft . , Nel . : 1 m . DGB , 1 5 . 2 . 66 , FRI, J . S . Dugdale .
REMARKS : Every series of Crambus dicrenellus examined by me in the New
When the specimens are in good condition t he two species can be distinguished at sight with a little experience . The forewing colour of is a
rich lustrous bronze-yellow intermediate between the colour of
and 0 . heteraulus . The termen of the forewings in is slightly
sinua.te , while that of O .dicrenellus is moderately convex , and the forewing
c olour is dull glossy brown.
Vfuile the female of O . di crenellus has yet to be collected, the males
of dicrenellus and are quite distinct on genitalia characters ,
having a thorn at the dorsal apex of the aedoeagus . The drawing_
of the male genitalia by Philpott ( 1 929 , fig. 35 ) , labelled Crambus dicrenellus Meyr . is based on a specimen of
The distribution of specimens of O .dicrenellus is coincidental with
the range of Chionochloa australis
(
carpet grass ) shown by Burrows ( 1 967) ,but
(
midribbed snowgrass) , Ch . rubra(
red tussock) and(
narrow-leaved snowgrass)
are also possibilitie s .In the phylogenetic scheme this species i s placed closer to O . flexuosellus and O . horistes than any other spe cie s .
Orocrambus flexuosellus
(
Doubleday) comb . nov .Crambus flexuo sellus Doubleday, 1 843 , in Dieffenbach ' s Travels in New Zealand, 2 , P • 289 .
Syntype m. in British Museum
(
Natural History)
, labelled "New Zealand42 - 55" .
EXTERNAL CHARACTERS(
Slide 6 ) : Detailed description by Meyrick ( 1 883 ) ,coloured illustration by Hudson ( 1 928, Plate XX fig . 31 ) and abbreviated
description, photograph by Gaskin ( 1 966 , Plate 1 8 , fig . 1 ) . Female frenulum
triple , eye nocturnal type . Easily separated from all other members of the genus so far known by the dark costa of the forewing and the dark subterminal markings interrupting the white forewing fascia . Wing span 1 9 - 26 mm.
(
bothsexe s
)
.MALE GENITALIA
(
Fig. 47 ) : As for generi c and species group except for following : Arms of vinculum about t uncus ; saccus tapered; co sta of valve with flange ,dorsal margin about
i
uncus . Sacculus with very small flange . Gnathostapered, pointed , equal to uncus . Aedoeagus 3-3-.lx uncus , length to median
breadth ratio 1 0 : 1 , dorsal line strong from apex to t, with subapical thorn dorsad .
FE��LE GENITALIA ( Fig . 1 34) : Eighth abdominal tergite fusing dorsa-laterally
with -ostium bursae , half as wide again at fusion as in dorsal midline . Eighth
sternite a weak lunate plate . Ductus bursae about 8x length of anal papillae ,
chitinised to �' with reverse loop at
�'
ductus seminalis joining at�.
Ostiumbursae narrowed at mouth, with exaggerated dorsal lip , ostiolar pouch swollen,
general appearance of ostiolar region vase-shaped ( urceolate ) . with two oval signa nearly equal in size .
Corpus bursae
EGG: Figured by Hudson ( 1 928, Frontispie ce , fig .
�) .
Oblate , 1 1 - 1 7 vertical ribs, 0.40 - 0.45 x 0 . 28 - 0.32 mm, creamy yellow when first laid, turning pinkish about 24 - 35 hours after being laid and vermillion after 4 - 7 days . First de scribed by Watt ( 1 914) . Duration of egg stage 9 - 20 days between October and February, 24 - 29 days in July, ( not ��er controlled conditions ) . Infertile eggs do not undergo the above colour changes . Aboutone third of the way through the period of development the orange colour deepens
to one side near the waist of the egg . This is the embryo becoming large and
distinct enough to be visible through the shell . About half way through the
development period the embryo become s detached from the wall of the egg except
in one small lateral area . This may approximate to the st�ges F-G shown for
the dragonfly by Imms ( 1 946 , after Ti[yard and Brandt ) , but the
embryology of O . flexuosellus has not been studied in detail in this work. After two thirds of the development of O .flexuosellus through the egg stage is complete , two approximated dark brown patches become visible . These are the lateral halves of the head capsule . Shortly before hatching a third can be seen, this is the pigmented prothoracic shield . At about this stage the last of the pigmented yolk is absorbed by the larva , leaving only some transparent fluid . This is taken up about twelve hours before emergence .
HATCHING BEHAVIOUR : Some movement of the larva may be seen up to two days b efore emergenc e , but in the last six or so hours movement of the he ad is
almost continuous . In 1 8 batche s
(
each containing 2 7 - 65 fertile eggs)
of this spe cie s maintained under unc ontrolled conditions , all hatching took place between 0700 hours and 1 200 hours , with 1 3 batche s completing their emergencebefore (800. In 3 batche s some hatching which did not occur on the first day
t ook place in the same period on the following day . In no batch did hatching
take place over a longer period than two days . The larva opens a hole in the
wall of the shell with a diagonal slicing movement of the mandible s . This hole
is usually on the " shoulder" of the curve of the oblate egg , but may occasionally